1986, Graduation from the Moscow Institute of Physics and Technology, USSR; 1989, PhD in Biology (membrane biophysics) in the A. A. Bogomoletz Institute of Physiology, Kiev, USSR; 2000, Habilitation in Physiology, Institute of Physiology, Faculty of Medicine, University of Giessen, Germany; 2000, Principal Investigator, IBMC, Porto, Portugal; 2010, Group Leader, IBMC, Porto, Portugal